Tar Heel/Coachman Formation

Tar Heel/Coachman Formation
Stratigraphic range: Early-Middle Campanian
~83.2–77.5 Ma
TypeFormation
Unit ofBlack Creek Group
UnderliesBladen Formation
OverliesMiddendorf Formation
Lithology
PrimarySiltstone, sandstone
Location
Coordinates34°36′N 78°30′W / 34.6°N 78.5°W / 34.6; -78.5
Approximate paleocoordinates35°00′N 50°12′W / 35.0°N 50.2°W / 35.0; -50.2
RegionNorth Carolina, South Carolina
CountryUnited States
Type section
Named forTar Heel, North Carolina

The Tar Heel Formation, also known as the Coachman Formation in South Carolina, is a Late Cretaceous (early to middle Campanian-aged) geologic formation in North Carolina and South Carolina, United States. It preserves fossils, including amber dating back to the Cretaceous period. A locality known as Phoebus Landing, has been dated to 78.5-77.1 Ma, and the formation has been overall dated to the early Campanian based on fossil pollen.[1][2][3]

Likely deposited in a nearshore coastal environment representing a lower shoreface, it contains a high diversity of vertebrate remains. It has one of the most diverse dinosaur faunas known from the former landmass of Appalachia, the majority of which are known from two sites: Phoebus Landing along the Cape Fear River in Bladen County, North Carolina, as well as Stokes Quarry in Darlington County, South Carolina.[4][5][6]

Fossil pollen grains suggest a subtropical to warm, moist temperate climate for the region, with an ecosystem largely dominated by flowering plants.[3] The Tar Heel/Coachman Formation appears to be roughly concurrent with the Marshalltown Formation of New Jersey, which preserves a similar fauna.[4]

Paleobiota

Cartilaginous fish

Based on the Paleobiology Database & Robb (1989):[7][8]

Cartilaginous fish of the Tar Heel/Coachman Formation
GenusSpeciesLocationNotesImages
"Asteracanthus"A. sp.Phoebus Landing, NCA hybodontid shark.
BrachyrhizodusB. wichitaensisPhoebus Landing, NCAn eagle ray.
CarchariasC. holmdelensisPhoebus Landing, NCA sand shark.
C. samhammeri
"Hypolophus"H. sp.Phoebus Landing, NCA whiptail stingray.
MeristodonoidesM. montanensisPhoebus Landing, NCA hybodontid shark.
RhombodusR. laevisPhoebus Landing, NCA rhombodontid ray.

Ischyrhiza

I. miraPhoebus Landing, NCA sawskate.
IschyodusI. cf. bifurcatusPhoebus Landing, NCA callorhinchid chimaera.
ScapanorhynchusS. texanusPhoebus Landing, NCA goblin shark.

Squalicorax

S. kaupiPhoebus Landing, NC

Stokes Quarry, SC

An anacorid shark. Includes a coprolite potentially assignable to S. kaupi, containing the vertebrae of a baby turtle.[9]
S. pristodontus
SquatinaS. hasseiPhoebus Landing, NCAn angelshark.

Ray-finned fish

Based on the Paleobiology Database, Robb (1989) & Stringer et al (2018). Some species are known only from otolith remains from Blue Banks & Auger Hole Landings, North Carolina, which are indicated:[7][8][10]

GenusSpeciesLocationMaterialNotesImages
Albula?A. campanianaBlue Banks & Auger Hole Landings, NC263-378 otolithsA bonefish.
?A. cf. A. ripleyensisBlue Banks Landing, NC1 otolith
A. sp.Phoebus Landing, NC
AnomoeodusA. phaseolusPhoebus Landing, NCA pycnodontid pycnodont.
Ariidae indet.Blue Banks Landing, NC11 otolithsA sea catfish of uncertain affinities.
Aulopidae indet.Blue Banks Landing, NC1 otolithA flagfin of uncertain affinities.
Beryx?B. maastrichtiensisBlue Banks & Auger Hole Landings, NC55 otolithsA potential relative of alfonsinos.
?B. zideki41 otoliths
?Congridae indet.Auger Hole Landing, NC1 otolithAn apparent conger eel of uncertain affinities.
CylindracanthusC. ornatusPhoebus Landing, NCA fish of uncertain affinities.
EnchodusE. cf. petrosusPhoebus Landing, NCAn enchodontid aulopiform.
Gonostomatidae indet.Blue Banks & Auger Hole Landings, NC4 otolithsA bristlemouth of uncertain affinities.
Hoplostethus?H. coffeesandensisBlue Banks & Auger Hole Landings, NC4 otolithsA slimehead.
KokenichthysK. ensisBlue Banks Landing, NC2 otolithsAn elopiform or osteoglossiform.[11]
Lepisosteidae indet.Phoebus Landing, NCA gar.
Megalopidae indet.Blue Banks Landing, NC1 otolithA tarpon of uncertain affinities.
OsmeroidesO. weileriBlue Banks Landing, NC6 otolithsAn osmeroidid elopomorph.
ParalbulaP. caseiPhoebus Landing, NCA phyllodontid elopomorph.
?Pempheris?P. huddlestoniAuger Hole Landing, NC1 otolithAn apparent sweeper.
Percoidei indet.Blue Banks & Auger Hole Landings, NC5 otolithsA percoid.
?Polymixia?P. cf. harderiBlue Banks Landing, NCAn apparent beardfish.
PterothrissusP. carolinensisBlue Banks Landing, NC67 otolithsA pterothrissine bonefish.
SaurodonS. sp.Phoebus Landing, NCA saurodontid ichthyodectiform.
StephanodusS. sp.Phoebus Landing, NCA pycnodont.
XiphactinusX. audaxPhoebus Landing, North Carolina, NCAn ichthyodectid ichthyodectiform.
X. vetus

Reptiles

Records from SC based on Schwimmer et al (2015):[6]

Dinosaurs

Ornithischians

Based mainly on Brownstein (2018):[4]

Ornithischians of the Tar Heel/Coachman Formation
GenusSpeciesLocationNotesImages

cf. Hadrosaurus

?H. foulkiiPhoebus Landing & Sampson County, NC[12]A hadrosaurid, tentatively assigned to the well-known H. foulkii.
"Hadrosaurus""H." minorPhoebus Landing, NCA small or juvenile hadrosaur, nomen dubium.

Hypsibema

H. crassicaudaPhoebus Landing & Sampson County, NCA gigantic hadrosauroid. First discovered in North Carolina from Sampson County, 1869, and described by Edward Drinker Cope. It was measured to be 12-17 meters, making it one of the largest hadrosauroids.[13] Type locality for genus and species.
Leptoceratopsidae indet.Sampson County, NCA leptoceratopsid ceratopsian. Known from an indeterminate left maxilla found in 2016.[14]

Lophorhothon

L. atopusPhoebus Landing, NCA hadrosauromorph. Originally described from the Mooreville Chalk Formation, Alabama.

An indeterminate hadrosauroid is known from Stokes Quarry, SC.[4]

Theropods
Theropods of the Tar Heel/Coachman Formation
GenusSpeciesLocationNotesImages
AppalachiosaurusA. montgomeriensisStokes Quarry, SCA large eutyrannosaur.[4]
cf. Coelosaurus (="Ornithomimus")C. sp.Phoebus Landing, NCAn intermediate ornithomimosaur.[4][15]
Dromaeosauridae indet.Sampson County, NCA large dromaeosaurid, larger than Saurornitholestes but smaller than Dakotaraptor.[16]
DryptosaurusD. sp.Phoebus Landing, NCA large dryptosaurid eutyrannosaur. Originally known from the Maastrichtian of New Jersey, but similar remains referable to this species have been recovered from the Tar Heel.[4] However, most of the attributed specimens are considered as indeterminate Eutyrannosauria separate from this genus, with only a partial femur fragment tentatively classified as D. sp.[17]

Saurornitholestes

S. langstoniBurches Ferry & Stokes Quarry, SC

A widespread dromaeosaurid ("raptor"). Specimens from this formation provided the first evidence of this species from Appalachia.[4][18][19]

Indeterminate theropods, ornithomimosaurs, and maniraptorans are known from Stokes Quarry.[4]

Crocodylomorphs

Based on the Paleobiology Database:[7]

GenusSpeciesLocationNotesImages
BorealosuchusB. formidabilisPhoebus Landing, NCA eusuchian.
BottosaurusB. sp.Stokes Quarry, SCAn early caiman.
DeinosuchusD. rugosus (=Thecachampsa rugosa, Polydectes biturgidus, Polyptychodon rugosus)Phoebus Landing & Clifton Farm, NC

Stokes Quarry, SC

An alligatoroid, one of the largest known crocodilians. Type locality for genus and species.
Gavialoidea indet.Stokes Quarry, SCA gavialoid.

Turtles

Based on the Paleobiology Database:[7]

GenusSpeciesLocationNotesImages
AdocusA. beatusPhoebus Landing, NC

Stokes Quarry, SC

An adocid.
BothremysB. cookiPhoebus Landing, NCA bothremydid side-necked turtle.
ChedighaiiC. barberiPhoebus Landing, NCA bothremydid side-necked turtle.
C. hutchinsoni
CorsochelysC. bentlyiStokes Quarry, SCA sea turtle, likely a dermochelyid. Type locality for species.
EuclastesE. wielandiStokes Quarry, SCA pancheloniid sea turtle.
OsteopygisO. emarginatusPhoebus Landing, NCA macrobaenid.
?Taphrosphys?T. daresPhoebus Landing, NCA bothremydid side-necked turtle, taxonomic placement disputed.
ToxochelysT. sp.Stokes Quarry, SCA toxochelyid sea turtle.
"Trionyx""T." halophilusPhoebus Landing, NCA stem-softshell turtle, likely not an actual member of the genus Trionyx. Species names are nomina dubia.
"T." priscusStokes Quarry, SC

Plesiosaurs

GenusSpeciesLocationNotesImages
Elasmosauridae indet.Stokes Quarry, South CarolinaAn elasmosaurid.

Squamates

GenusSpeciesLocationNotesImages
HalisaurusH. sp.Phoebus Landing, NCA halisaurine mosasaur.
PlatecarpusP. sp.Phoebus Landing, NCA plioplatecarpine mosasaur.
PrognathodonP. sp.Phoebus Landing, NCA mosasaurine mosasaur.
Teiidae indet.Stokes Quarry, SCA teiid lizard.
TylosaurusT. sp.Phoebus Landing, NC

Stokes Quarry, SC

A tylosaurine mosasaur.

Mammals

Based on the Paleobiology Database:[20]

GenusSpeciesLocationNotesImages
Marsupialia indet.Darlington County, SCA presumed marsupial relative.[20]
Multituberculata indet.Darlington County, SCA multituberculate.[20]

References

  1. ^ Longrich, Nicholas R. (2016). "A ceratopsian dinosaur from the Late Cretaceous of eastern North America, and implications for dinosaur biogeography". Cretaceous Research. 57: 199–207. Bibcode:2016CrRes..57..199L. doi:10.1016/j.cretres.2015.08.004.
  2. ^ "Geolex — TarHeel publications". ngmdb.usgs.gov. Retrieved 2024-11-13.
  3. ^ a b Mitra, Madhumi (2002-07-29). Paleopalynology of the Tar Heel Formation of Atlantic Coastal Plain of North Carolina, United States (Thesis).
  4. ^ a b c d e f g h i Brownstein, Chase D. (2018-02-08). "The biogeography and ecology of the Cretaceous non-avian dinosaurs of Appalachia". Palaeontologia Electronica. 21 (1): 1–56. doi:10.26879/801. ISSN 1094-8074.
  5. ^ "PBDB Collection". paleobiodb.org. Retrieved 2024-11-13.
  6. ^ a b Schwimmer, David R.; Sanders, Albert E.; Erickson, Bruce R.; Weems, Robert E. (2015). "A Late Cretaceous Dinosaur and Reptile Assemblage from South Carolina, Usa". Transactions of the American Philosophical Society. 105 (2): i–157. ISSN 0065-9746. JSTOR 24398224.
  7. ^ a b c d Czaplewski, John J. "PBDB Navigator". paleobiodb.org. Retrieved 2024-10-10.
  8. ^ a b Albert J Robb III (1989). The Upper Cretaceous (Campanian, Black Creek Formation) fossil fish fauna of Phoebus Landing, Bladen County, North Carolina.
  9. ^ Schwimmer, David R.; Weems, Robert E.; Sanders, Albert E. (2015). "A Late Cretaceous Shark Coprolite with Baby Freshwater Turtle Vertebrae Inclusions". PALAIOS. 30 (9/10): 707–713. Bibcode:2015Palai..30..707S. doi:10.2110/palo.2015.019. ISSN 0883-1351. JSTOR 44708722.
  10. ^ Stringer, Gary L.; Clements, Don; Sadorf, Eric; Shannon, Kevin (2019). "First Description and Significance of Cretaceous Teleostean Otoliths (Tar Heel Formation, Campanian) from North Carolina". Eastern Paleontologist. 4: 1–22. ISSN 2475-5117.
  11. ^ Stringer, Gary L.; Sloan, James Carson (2023-05-25). "First Cretaceous teleostean otolith assemblage (Arkadelphia Formation, upper Maastrichtian) from Arkansas, USA, early Gadiformes, and the Western Interior Seaway". PaleoBios. 40 (3). doi:10.5070/P940361192. ISSN 2373-8189.
  12. ^ NCSM Paleontology Database
  13. ^ Miller, Halsey W. (1967). "Cretaceous Vertebrates from Phoebus Landing, North Carolina". Proceedings of the Academy of Natural Sciences of Philadelphia. 119: 219–239. ISSN 0097-3157. JSTOR 4064614.
  14. ^ Longrich, Nicholas R. (2016-01-01). "A ceratopsian dinosaur from the Late Cretaceous of eastern North America, and implications for dinosaur biogeography". Cretaceous Research. 57: 199–207. Bibcode:2016CrRes..57..199L. doi:10.1016/j.cretres.2015.08.004. ISSN 0195-6671.
  15. ^ Baird D., and Horner, J., 1979, "Cretaceous dinosaurs of North Carolina" Brimleyana 2: 1–28
  16. ^ Brownstein, Chase D. (2018-12-01). "A large dromaeosaurid from North Carolina". Cretaceous Research. 92: 1–7. Bibcode:2018CrRes..92....1B. doi:10.1016/j.cretres.2018.07.006. ISSN 0195-6671. S2CID 135459468.
  17. ^ Mortimer, Mickey (2024). "Tyrannosauroidea". The Theropod Database. Retrieved Jan 28, 2025.
  18. ^ Brownstein, Chase D. (2018-04-05). "Diversity of raptor dinosaurs in southeastern North America revealed by the first definite record from North Carolina". PeerJ Preprints. doi:10.7287/peerj.preprints.26829v1.
  19. ^ Brownstein, Chase D. (2018). "The distinctive theropod assemblage of the Ellisdale site of New Jersey and its implications for North American dinosaur ecology and evolution during the Cretaceous". Journal of Paleontology. 92 (6): 1115–1129. Bibcode:2018JPal...92.1115B. doi:10.1017/jpa.2018.42. ISSN 0022-3360.
  20. ^ a b c "PBDB Collection". paleobiodb.org. Retrieved 2024-11-15.
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